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The game takes place in a fictional floating world called Cocoon, whose government, Sanctum, sends its citizens down for suspected connections to Pulse, a large, frightening world below. A female soldier codenamed Lightning defies the authorities and fights them to save her sister, chosen by the gods of Pulse and an enemy of Cocoon. Soon, other characters join her, and all of them become chosen to fulfill some unknown purpose. The heroes confront Sanctum, trying to figure out their purpose; if they succeed, they will turn into crystals and gain eternal life; if they fail, they will become terrible monsters.